David Denton, Jim Hamilton, Matt Scott and Dougie Fife will all start for Scotland against England on Saturday after Vern Cotter made five changes to his starting XV.
Stand-off Finn Russell also returns after suspension ruled him out of the 22-19 defeat to Italy at Murrayfield two weeks ago.
Edinburgh No.8 Denton – the only player among the incoming five not to have featured in this year’s competition – replaces Johnnie Beattie after shaking off the calf strain which has kept him out of action so far.
Fife and Scott replace injured wing Sean Lamont and centre Alex Dunbar while Russell starts in place of Peter Horne and experienced Saracens lock Hamilton takes the place of Tim Swinson.
Cotter said: “Jim comes in and will bring his physicality and understanding of English rugby to our forward pack.
“It’s good to have Dave Denton back as he provides us with strong ball-carrying and strong defence. He’s also a very good lineout forward and will give us a bit more physical density against a big forward pack.
“It’s been tough on Finn to sit out and it’s great to have him back. He’s slotted straight back in to the structure we’re looking for.
“Finally, Dougie came on and played well against France and has had a couple of games with his club.
“We’re looking for him to bring his enthusiasm, energy and high work-rate, particularly in kick-chase and receipt.”
“This is a very important game. It will be played away from home at a very intense level and will allow us to assess further our ability to operate away from home in a hostile environment.
“Our focus, however, has been on ourselves and how we can perform better, by identifying the areas that we can control – like improving our skill sets and reinforcing our cohesion – to withstand the difficult times and also apply some pressure.”
Scotland: Stuart Hogg; Dougie Fife, Mark Bennett, Matt Scott, Tommy Seymour; Finn Russell, Greig Laidlaw (Capt.); Alasdair Dickinson, Ross Ford, Euan Murray; Jim Hamilton, Jonny Gray; Robert Harley, Blair Cowan, David Denton .
Replacements: Fraser Brown, Ryan Grant, Geoff Cross, Tim Swinson, Johnnie Beattie, Adam Ashe, Sam Hidalgo-Clyne, Greig Tonks.
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